Plant Biology - Roots

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Study GuidePlant BiologyRoots1. Primary Root Tissues and StructureTheprimary roothas a simpler tissue arrangement than theprimary stem. This is because roots donot produce leaves, so they do not need side connections to the vascular system. As a result, theirinternal structure is more straightforward.Theprimary plant bodyof the root is formed by three primary meristems and has three mainregions:1.Epidermisthe outer protective layer2.Cortexa broad middle region mainly for storage3.Stele (vascular cylinder)the central conducting region1.1EpidermisTheepidermisis the outermost layer of the root. In young roots, many epidermal cells formroothairs.Why root hairs are important:They greatlyincrease surface areaThis allows morewater and mineral absorptionEach root hair is an extension of theprotoplast(living cell content)Root hairs:Havevery thin wallsAreextremely delicateand easily brokenLive only for a short timeIn some plants, the root epidermis is covered by athin waxy cuticle. This layer is not thick enough toblock the movement of water and dissolved substances into the root.

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Study Guide1.2CortexThecortexlies between the epidermis and the stele and makes up thelargest portionof the primaryroot. It is composed mainly ofparenchyma cells.In manydicots (eudicots)andgymnospermsthat later develop extensive secondary growth, thecortex is eventuallycrushed, and its storage role is taken over by other tissues.The cortex hasthree distinct layers:1.Hypodermis (exodermis)2.Storage parenchyma3.Endodermis1.3Hypodermis (Exodermis)Thehypodermislies just beneath the epidermis and is usuallyone or two cell layers thick.Key features:Cell walls areheavily suberizedWalls areimpermeable to waterCells aretightly packedwith no air spacesFunctions:Preventsloss of water and nutrientsfrom the cortexProtects againstsoil microorganismsEspecially well developed inarid-region plantsand plants withshallow roots1.4EndodermisTheendodermisis theinnermost layer of the cortexand is easy to recognize because of specialstructures calledCasparian strips.Casparian strips:Are bands ofsuberinOccur on theradial and transverse wallsBlock movement through cell walls

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Study GuideWhy this matters:Water and dissolved minerals must passthrough living cell membranesThis allowsselective absorptionHarmful substances in the soil are prevented from entering the plantThe endodermis is:Almost always present in rootsRarely found in stems1.5Storage ParenchymaThestorage parenchymamakes up most of the cortex.Characteristics:Thin-walled, living cellsLoosely packed withmany air spacesStorestarch and other substancesFunctions:Food storageProvidesaeration, allowing roots to get oxygenCells expand and shrink as materials move in and out1.6Stele (Vascular Cylinder)Thesteleincludes all tissuesinside the cortex, namely:PericycleXylemPhloemPith(in some plants)In mostdicot roots, the center is filled withsolid xylem.In mostmonocot roots, the center contains aparenchymatous pith.

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Study Guide1.7PericycleThepericycleis a thin layer of parenchyma cells located justinside the endodermis.Important roles:Cellsretain the ability to divideGives rise tolateral (branch) rootsDuring secondary growth, it forms:oVascular cambiumoOften thefirst cork cambiumThe pericycle can also produce more pericycle cells through division.1.8Vascular TissuesThe arrangement ofxylem and phloemdiffers between plant groups:Dicot (eudicot) rootsoCentral, lobedxylem coreoPhloemlocated between xylem armsoThis arrangement is called aprotosteleMonocot rootsoXylem forms a ringoSurrounds a centralpithoThis arrangement is called asiphonosteleThese vascular patterns are important because they help scientiststrace evolutionary relationshipsamong plants.

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Study GuideSummaryRoots have asimpler structurethan stemsThe cortex is mainly forstorage and regulationThe endodermis ensuresselective absorptionThe pericycle formslateral rootsXylem and phloem arrangement differs in dicots and monocotsThis organized structure allows roots toabsorb, store, regulate, and transportmaterials efficientlywhile anchoring the plant firmly in the soil2. Secondary Growth of Roots2.1What Is Secondary Growth in Roots?Secondary growth is the process thatincreases the thickness (girth)of roots.It is especially important inwoody perennial plants, where secondary tissues make upmost of theroot mass.Primary tissuescontinue to form only in young feeder roots.Themain supporting root systemis made ofsecondary tissues.These tissues are produced bylateral meristems:oVascular cambiumoOne or more cork cambiaAlthough roots grow underground and are not easily seen, they areas massive as the aerial partsof trees.They balance the weight of the plant above the ground andkeep the tree upright and stable.
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